Mary Keene is placed in John Ingalls' office to help carry out a plot to steal his money by placing a fake sister in his home. Mary, however, falls in love with Ingalls, foiling the scheme.

George Woods is leading a happy and contented life with his wife and little daughter in the small town of Milford when millionaire Cyrus Morton from New York offers to make him his chauffeur. George accepts and goes to the city where he finds everyone preoccupied with the pursuit of wealth. The fever enters his blood, and upon finding Morton's lost wallet, George takes the money and plunges into the stock market. His investment is successful, and from then on he becomes obsessed with money, forgetting his wife and child in his vain pursuit. But even with the vast wealth he accumulates, George finds that he is unhappy. Through his wife Mary's efforts, he finally realizes that true happiness cannot be found in the accumulation of riches, but lies in simpler pleasures.

A baby, found strapped to a donkey in the North Carolina mountains where her father drowned, is called Twilight and raised by the Anwells. At sixteen, Twilight, in love with her foster brother Jim, who runs the family's lumber business, grows jealous when Elise Charmant, vacationing with her father, a brain specialist, monopolizes Jim at the Fireman's Ball.