Four years after the events of SHIROBAKO the series, Musashino Animation has gone through unprecedented changes and things are looking bleak for the studio. One day, Aoi Miyamori is offered a role managing a new theatrical animation project. With only a few months left before the deadline, can MusAni complete the project in its current state? It’s up to Aoi to work alongside both familiar faces and new to finish the project in time!

14-year-old Nozomi Onda has only one thing on her mind: playing beautiful soccer. There's just one problem: no matter how much she longs to participate in official matches, she'd have physically superior boys as opponents. But when a boy from her past confronts her on the street, she decides she can't wait any longer.