Lupin has invited all of his friends to a villa on a private island. It's the first time they've seen each other for the past five years. Lupin is planning a big surprise while they are all at the villa except Fujiko, Goemon and Jigen find Lupin's behavior to be unusual. They are also curious about who the man in the iron mask is, who is chained up to a wall for the reasons they are about to find out.

While performing a puppet show at a nursery, the HappinessCharge Pretty Cures are surprised when one of the dolls starts talking. “Save our kingdom!” Tsumugi, the ballerina doll, pleads. The Pretty Cures accompany Tsumugi to the land of dolls, a girls’ paradise of cute stuffed animals, talking dolls, music and dance ruled by the noble Prince Zeke. But the kingdom is under attack by the forces of Black Fang, a mysterious man who wants to conquer the land for his own nefarious purposes. As sorrow spreads throughout the Kingdom of Dolls, the Pretty Cures come to the rescue.

Space Pirate Captain Harlock and his fearless crew face off against the space invaders who seek to conquer the planet Earth.