When brilliant video game maker Flynn hacks the mainframe of his ex-employer, he is beamed inside an astonishing digital world and becomes part of the very game he is designing. In his mission through cyberspace, Flynn matches wits with a maniacal Master Control Program and teams up with Tron, a security measure created to bring balance to the digital environment.

In honor of his birthday, San Francisco banker Nicholas Van Orton, a financial genius and a cold-hearted loner, receives an unusual present from his younger brother, Conrad: a gift certificate to play a unique kind of game. In nary a nanosecond, Nicholas finds himself consumed by a dangerous set of ever-changing rules, unable to distinguish where the charade ends and reality begins.