Bob Parr has given up his superhero days to log in time as an insurance adjuster and raise his three children with his formerly heroic wife in suburbia. But when he receives a mysterious assignment, it's time to get back into costume.

Remy, a rat, possesses a palate far more refined than that of his fellow comrades. He dreams of becoming a chef, one who creates rather than scavenges. When fate deposits him in the sewers beneath one of Paris’s most famous restaurants, he finds himself ideally placed to fulfill his dream. Forming an unusual alliance with a hapless young kitchen worker, Remy begins a daring culinary double life. As Remy pursues his vision, he must navigate the suspicions of the calculating Head Chef Skinner, the disapproval of Remy’s own colony, and the foreboding presence of renowned food critic Anton Ego, who strikes fear in the hearts of chefs all throughout France.