Blamed for the theft of an orphans fund, Captain James Wynnegate flees to the West where he makes a new life with the Indian woman Nat-U-Rich.

The one was a venturer - the other an adventurer - the one a man who wanted to see adventure, but who had never been beyond the city limits - the other a man who had seen adventure in all parts of the world, and who assured the venturer that things were just as monotonous every place in the world as in the city. So they met, each seeking for the unconventional, on a New York street, and dined together as men out of luck, with two cents between them - and still nothing happened. They both had credit at the hotel. Then into their lives came the feminine influence - a sweet girl who lived in a house which was irrevocably a household. The adventurer hesitated - he had yet to satisfy his longing for the incalculable. Suddenly love changed the venturer into an adventurer, and settled the adventurer into a venturer.

At a reception given for the vacationing Prince Zarl of Zorania, secretly the emissary of Zorania set on negotiating a treaty with the United States, Geoffrey Wynne, apparently a society dandy, but in reality a secret service agent, meets the prince. During the reception, Wynne is summoned to Washington where he learns that the treaty has been stolen and is being held for $15,000,000 ransom. Discovering that one of the thieves is an Italian named Farnelli, Wynne enlists the aid of his fiancée, Irene Mitchell, in apprehending the thief. Irene meets the Italian who offers to accompany her to New York. En route, Wynne overtakes them, chloroforms Farnelli and rips off his disguise to reveal Prince Zarl. Zarl then admits that he has stolen the treaty in order to cover his gambling debts.

Oma loves Jack and refuses to marry Hartman. Jack protects her and they eventually get married. Lafe and Hazelet have been good friends for years. Hartman tells Lafe that Hazelet voted dry. Lafe and Hazelet have a fight. Lafe builds a spite fence so Oma and Jack cannot communicate. Lafe dies with delirium tremens. Mrs. Tuthill dies from injuries received from Lafe while he is drunk. Just before Mrs. Hazelet passes away her husband promises her never to vote wet again. Maude Le Fever, the siren, who married Alfred Tuthill, deserts him and runs away with Sam Weber; she later deserts him. Alcohol kills old Dan Weber and leaves Sam a poverty-stricken wretch. Alfred Tuthill is redeemed from rum and builds a picture theater where the old saloon once stood and becomes a respected citizen. Jack is elected Mayor of the city, defeating Hartman by a large majority.

Hal Page, the weakling brother of Stephen Page, the town mayor, falls in love with Carline Shrefton, who throws over Burt Staley to entangle Hal in her self-serving schemes. Furious over her abrupt departure, Staley shows up at Carline's, and a jealous fight with Hal ensues in which Staley is shot and killed with Carline's gun. After Hal confesses to the deed, Stephen quietly sends him off to Spain. One year later, Stephen announces his engagement to Marion Hayward, the daughter of the district attorney, who is pressing charges against James Reed, a corrupt politician and Carline's new husband. To save Reed, Carline threatens Stephen with exposure, but he refuses to bend to her demands. Just days before Carline is to reveal her story about Staley, Hal appears and confesses to Hayward that Carline is the true killer.

The story centers on Flapjack Ike (played by John Bunny), a dominant local figure or "autocrat" in the titular town. Ike faces off against Edith, the Widow (played by Flora Finch), who challenges his authority.

Polly Biggs helps care for her younger brother and sisters, while her widowed mother works hard as a seamstress to earn a living. Mayor Hoadley, Mrs. Biggs' brother, a crooked politician, calls to sympathize with her on the death of her husband, and Polly takes a dislike to him.