An innocent rubbish tip assistant, Gert, is given the task of escorting a beautiful mute "princess", Plum, on a perilous journey to meet her prospective husband. Their journey is fraught with danger, challenges, romance and discovery. In surreal action, they encounter a menagerie of characters and animals in far-flung corners of New Zealand.

Mater the tow truck travels from country to country as he retells his infamous but unbelievable stories.

A routine tow lands Mater in Tokyo, where he is challenged to a drift-style race against a nefarious gang leader and his posse of ninjas. With the help of his friend, 'Dragon' Lightning McQueen, and some special modifications, Mater attempts to drift to victory and become Tow-ke-O Mater, King of all Drifters.

With the help of an autotuning AI, a pizza delivery Gal plays a deadly initiation game to join the punk hacker gang who killed her father.