Young, innocent, confiding, it is a shock to Ann Fenton to learn that her supposed husband is not a business man, but a gambler, and that her marriage is bigamous. The child is taken from her by a Helping Hand Society and apprenticed to a brutal farmer. She is left upon her own resources. Seven years later Fenton again crosses her path, but she finds happiness in honorable marriage while her betrayer is taken away to face a murder charge, and the Song of the Soul now rises in full, pure tones from the breast of the happy wife and mother.

Dishonored by playboy John Radon, simple country girl Stella Dean flees to the city to hide her shame. Leading a disreputable life she eventually obtains great wealth as a courtesan known as the Black Nightingale. One day she meets Milton Taylor, an artist from her hometown who knew her when she was an innocent, and he asks her to pose as his model of the Madonna. Stella agrees and feels cleansed by the experience, however when Milton discovers Stella's reputation he begins to drink and leaves her, his illusions shattered. Repentant, Stella converts her mansion into a refuge for foundlings and returns to her hometown of Pleasantville reconciling with Milton.