Simon Cowell is shot at his birthday party, and Ant and Dec are accused and forced to run.

Pudsey the cheeky London stray dog is quite happy being a lone ranger, looking out for number one, until he meets siblings Molly, George and Tommy. After losing their father, their mother Gail is moving the family to the sleepy village of Chuffington and Pudsey tags along, to the dismay of their landlord, Mr Thorne, and his cat Faustus. As Pudsey starts to settle in with the family and realises what he was missing when he was alone, he stumbles across Thorne's evil plan, and he determines to save them and the whole village.